A network typically counts clicks based on accesses to their click tracking redirect. These redirects are applied with your ad on the publisher page. Because of this, the network will typically get the first click look of any particular click. The network may then apply some filters to the final presented click count. For example Google Ads has an invalid click count metric that captures some GIVT that is typically not shown by default.
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Spider AF counts clicks on the final destination page, where it determins where the click comes from based on the URL and referral parameters of each session. Because of this Spider AF may not execute on all network counted clicks.
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Additionally there are cases where Spider AF may over count clicks. When sessions continue or are cached with the same URL parameters, for instance if a user clicked an ad, closed the browser, and reopened the browser later causing the page to load again, Spider AF may count additional clicks.
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For more information about how Spider AF counts clicks see here: How Spider AF Counts Clicks